The second son of Lord Gwyn in the game Dark Souls I. His affinity with the moon, a traditionally female trait, caused him to be raised as a girl.
Wears a spiked, sun-shaped mask with a veil, a long white robe, and has a lower body composed entirely of numerous tentacular snakes. Skilled in sorcery, illusions, and archery.
By the time of Dark Souls III, Gwyndolin has been consumed by Aldrich.